Breeding resident
The only breeding site for the Gannet in Wales is on Grassholm. Williams (1978) provides a résumé of the history of the colony which seems to have been established shortly before 1860, when 20 pairs were present, though Gurney (1913) suggests birds may have been present as early as 1820.
They were beset by problems during the early years, mainly from egg collectors, while in 1890 members of the Cardiff Naturalists Society, who were camping on the island at Whitsun, witnessed a party of naval and military officers from Pembroke Dock shooting Gannets and taking and smashing eggs. The miscreants were brought before the bench at Haverfordwest in August of that year and fined a total of £22.17s. in what the magistrates described as "a somewhat novel case ... but it was also a charge that could not be passed over lightly."
The growth of the colony aroused the interest of a succession of ornithologists, resulting in a series of nest counts, initially made by eye, then by photographs taken on the ground first by Acland and Salmon (1924), followed by the first aerial survey in 1956 (Lockley 1957); this remains the standard census method (see Table 1). Colony growth has not been uniform, Nelson (1978) demonstrating that Grassholm must have exported breeding birds in the 1960s but imported them in the 1920s and 1970s.
A fresh count of the Grassholm gannetry is clearly overdue as observations show a continued advance by the Gannets. The days are gone when visitors stood on the summit of the island viewing the colony spread below; the high point is now fully occupied by breeding birds which are advancing southwards across the island. The RSPB, who have owned Grassholm since 1948, have had to prohibit landing at two points in order to minimise disturbance to birds nesting in their vicinity, it being a matter of conjecture how long it will be before the sole remaining landing point is also occupied by breeding birds.
With apparently plenty of food to support their ever increasing numbers and few enemies the Grassholm Gannets have little to fear save for manmade hazards. The most obvious of these is that posed by entanglement in nylon lines and netting. The Gannets gather prodigious quantities of nesting material, normally seaweeds with the addition of grasses pulled from near the colony. However, anything floating in the sea which can be carried is likely to be incorporated into the nest. For the past two decades this has included nylon, and some nests can be bright orange in colour, so significant is the quantity. Adults and chicks can become entangled and deaths occur, either through birds being anchored to their nests or by becoming so entangled that they cannot fly properly.
Over 5,000 Gannets were ringed on Grassholm, largely by the staff of the Skokholm Bird Observatory, up until the late 1960s. There have been recoveries from as far apart as the Faeroes and Senegal, with most immatures following the traditional route south to west African waters where some remain for the first two years of life. Most adults do not travel so far but remain in home waters throughout the winter.
Large passages occur off Strumble Head in the autumn which probably include many birds from colonies other than Grassholm. Passage peaks in August, the average rate from 14 counts made in 1980 and 1981 being 180 birds per hour, the maximum 1,700 in five hours on 25 August 1980. An albino recorded at Strumble Head on 15 September 1985 was also seen passing Towan Head, Cornwall, on 16 September 1985, indicating through passage.
Although fairly sparsely distributed inshore during the winter months, typically less than ten at any locality, onshore gales temporarily push greater numbers towards the land; for example, up to 100 were seen at Strumble Head on 2 January 1988. Grassholm is usually deserted from about late October and reoccupied in January.
Fieldwork 1984-88
Red = breeding confirmed
Orange = breeding probable
Yellow = breeding possible
Donovan J.W. & Rees G.H (1994), Birds of Pembrokeshire